Having failed to advance past the group stage of Europe's elite club competition in their previous four attempts, Dinamo produced a morale-boosting performance in the Maksimir stadium.
Organised defending aided by a packed midfield frustrated a disjointed Arsenal and the home side always looked dangerous on the break, with Algeria striker El Hilal Arabi Soudani at the heart of all their moves.
Arsenal held the upper hand in the opening 20 minutes as Giroud missed two good chances, hitting the post with a scuffed effort after Dinamo's Portuguese keeper Eduardo had kept out the Frenchman's header with an acrobatic save.
Worse followed for the visitors when Giroud was shown a second yellow card for a reckless challenge after he was booked for dissent.
Sensing blood, a fired-up Dinamo came storming out after the interval and livewire Soudani, a handful all night for his markers, hit the post with a glancing header barely a minute into the second half.
The home fans were in raptures in the 58th minute when Chilean junior Fernandes rose at the near post to head home an inviting Paulo Machado corner past a static Arsenal defence.
Arsenal manager Arsene Wenger then replaced Oxlade-Chamberlain with Walcott and the winger made a quick impact, racing clear of two markers before he beat the advancing Eduardo with a sublime finish in the 79th minute.
Arsenal pressed forward in the closing stages but Dinamo held firm with some last-gasp interceptions in their penalty box.
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